01 - Preheat oven to 375°F. Line a 12-cup muffin tin with paper liners or grease lightly.
02 - In a large bowl, whisk together flour, poppy seeds, baking powder, baking soda, and salt. Set aside.
03 - In a separate bowl, rub sugar and lemon zest together with your fingers to release aromatic oils.
04 - Add eggs, Greek yogurt, vegetable oil, lemon juice, and vanilla extract to the sugar mixture. Whisk until smooth.
05 - Pour the wet ingredients into the dry ingredients bowl. Stir gently until just combined, avoiding overmixing.
06 - Spoon batter evenly into muffin cups, filling each approximately three-quarters full.
07 - Bake for 18 to 20 minutes, or until a toothpick inserted into the center comes out clean.
08 - Let muffins cool in the pan for 5 minutes before transferring to a wire rack to cool completely.